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At Firelink Shrine, on the roof that is above Petrus of Thorolund. To reach the roof, the player must ride the elevator leading to Undead Parish and jump off at the opening just above the Firelink Shrine entrance. After this, walk out on the right cliff and jump or roll onto the nearby pillar and walk into the building it is connected to, then the player can drop from there onto the roof to get the key.