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Found in the New Londo Ruins, after the water level is lowered.
Giving the Very Large Ember to Andre of Astora gives him the ability to ascend a standard weapon from +10 to +11, allowing for further standard weapon reinforcement from +11 to +15 (the highest it can go).
Taking the Very Large Ember to Rickert of Vinheim will cause him to comment on it, suggesting that the player take it to Andre of Astora.